Costly signals in a field cricket can indicate high- or low-quality direct benefits depending upon the environment.

Abstract excerpt

The handicap hypothesis proposes that male signals provide reliable information to females because only males of high condition provide high-quality mating benefits and can afford the costs of producing attractive signals. In the context of direct benefits, the handicap hypothesis predicts that benefit quality and signal attractiveness will positively covary among genotypes, positively covary among environments,...

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